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  <front>
    <journal-meta>
      <journal-id journal-id-type="publisher-id">17</journal-id>
      <journal-id journal-id-type="index">urn:lsid:arphahub.com:pub:8E638694-B4E0-570A-856A-746FF325BF6B</journal-id>
      <journal-id journal-id-type="aggregator">urn:lsid:zoobank.org:pub:FEF66878-15EE-4F8B-B369-7652D735020E</journal-id>
      <journal-title-group>
        <journal-title xml:lang="en">Research Ideas and Outcomes</journal-title>
        <abbrev-journal-title xml:lang="en">RIO</abbrev-journal-title>
      </journal-title-group>
      <issn pub-type="epub">2367-7163</issn>
      <publisher>
        <publisher-name>Pensoft Publishers</publisher-name>
      </publisher>
    </journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.3897/rio.12.e186726</article-id>
      <article-id pub-id-type="publisher-id">186726</article-id>
      <article-id pub-id-type="manuscript">30222</article-id>
      <article-categories>
        <subj-group subj-group-type="heading">
          <subject>Commentary</subject>
        </subj-group>
        <subj-group subj-group-type="scientific_subject">
          <subject>Life sciences</subject>
        </subj-group>
      </article-categories>
      <title-group>
        <article-title>The Final Ascent: from refuges to ruins in a warming world</article-title>
      </title-group>
      <contrib-group content-type="authors">
        <contrib contrib-type="author" corresp="yes">
          <name name-style="western">
            <surname>Wanke</surname>
            <given-names>Dominic</given-names>
          </name>
          <email xlink:type="simple">dominic.wanke@smns-bw.de</email>
          <uri content-type="orcid">https://orcid.org/0000-0001-5390-8993</uri>
          <xref ref-type="aff" rid="A1">1</xref>
          <xref ref-type="aff" rid="A2">2</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Whitmore</surname>
            <given-names>Daniel</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0000-0002-6051-5925</uri>
          <xref ref-type="aff" rid="A1">1</xref>
          <xref ref-type="aff" rid="A2">2</xref>
        </contrib>
      </contrib-group>
      <aff id="A1">
        <label>1</label>
        <addr-line content-type="verbatim">State Museum of Natural History, Rosenstein 1, D-70191, Stuttgart, Germany</addr-line>
        <institution>State Museum of Natural History</institution>
        <addr-line content-type="city">Rosenstein 1, D-70191, Stuttgart</addr-line>
        <country>Germany</country>
      </aff>
      <aff id="A2">
        <label>2</label>
        <addr-line content-type="verbatim">Center of Excellence for Biodiversity and Integrative Taxonomy (KomBioTa), Wollgrasweg 23, D-70599, Stuttgart, Germany</addr-line>
        <institution>Center of Excellence for Biodiversity and Integrative Taxonomy (KomBioTa)</institution>
        <addr-line content-type="city">Wollgrasweg 23, D-70599, Stuttgart</addr-line>
        <country>Germany</country>
      </aff>
      <author-notes>
        <fn fn-type="corresp">
          <p>Corresponding author: Dominic Wanke (<email xlink:type="simple">dominic.wanke@smns-bw.de</email>).</p>
        </fn>
        <fn fn-type="edited-by">
          <p>Academic editor: </p>
        </fn>
      </author-notes>
      <pub-date pub-type="collection">
        <year>2026</year>
      </pub-date>
      <pub-date pub-type="epub">
        <day>27</day>
        <month>02</month>
        <year>2026</year>
      </pub-date>
      <volume>12</volume>
      <elocation-id>e186726</elocation-id>
      <uri content-type="arpha" xlink:href="http://openbiodiv.net/01FC435B-CF87-5EF8-9E6F-72F2ADCF4139">01FC435B-CF87-5EF8-9E6F-72F2ADCF4139</uri>
      <uri content-type="zenodo_dep_id" xlink:href="https://zenodo.org/record/0">0</uri>
      <permissions>
        <copyright-statement>Dominic Wanke, Daniel Whitmore</copyright-statement>
        <license license-type="creative-commons-attribution" xlink:href="http://creativecommons.org/licenses/by/4.0/" xlink:type="simple">
          <license-p>This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.</license-p>
        </license>
      </permissions>
      <abstract>
        <label>Abstract</label>
        <p>Mountains exemplify how climate change transforms long-standing refuges into zones of loss. As species track suitable climates upslope, they confront absolute ecological limits. This commentary reflects on montane extinctions as a warning signal for broader planetary boundaries and the future of biodiversity in the Anthropocene.</p>
      </abstract>
      <kwd-group>
        <label>Keywords</label>
        <kwd>Anthropocene</kwd>
        <kwd>biodiversity</kwd>
        <kwd>climate change</kwd>
        <kwd>mountain ecosystems</kwd>
        <kwd>refugia</kwd>
      </kwd-group>
      <counts>
        <fig-count count="0"/>
        <table-count count="0"/>
        <ref-count count="7"/>
      </counts>
    </article-meta>
  </front>
  <body>
    <sec sec-type="Ascent to Extinction">
      <title>Ascent to Extinction</title>
      <p>Mountain ecosystems have long functioned as both cradles and repositories of biodiversity. Their topographic complexity, climatic heterogeneity and relative isolation have promoted speciation and served as safe havens during glacial cycles (<xref ref-type="bibr" rid="B13845955">Steinbauer et al. 2016</xref>). The mountainous areas of Southwest Asia, including Iran, represented key refugia during the Last Glacial Maximum (LGM), sustaining populations that later diversified into a rich mosaic of endemic species (<xref ref-type="bibr" rid="B13845891">Noroozi et al. 2018</xref>). These montane endemics often occupy narrow ecological niches finely tuned to local climatic conditions, making them particularly sensitive to environmental changes (<xref ref-type="bibr" rid="B13845939">Rahbek et al. 2019</xref>). Today, these same mountains illustrate a profound paradox. As global temperatures rise, montane species increasingly shift their ranges upslope to track favourable climatic conditions (<xref ref-type="bibr" rid="B13845902">Pauli et al. 2012</xref>; <xref ref-type="bibr" rid="B13845882">Freeman et al. 2018</xref>). Yet, the vertical space that once offered refuge now imposes an absolute limit, the mountain summit. This process, often described as the escalator to extinction, transforms ancient centres of survival into zones of irreversible loss (<xref ref-type="bibr" rid="B13845872">Colwell et al. 2008</xref>). The populations that have been pushed back to these ecological upper limits on mountain peaks are threatened with local extinction and, ultimately, total extinction, due to the lack of other suitable habitats.</p>
      <p>The story of montane endemics mirrors the broader trajectory of life on Earth in the Anthropocene. Just as mountain species are constrained by topography, humanity is confined by the ecological boundaries it has disrupted. The biosphere, our planetary refuge through countless epochs, now faces unprecedented pressures from climate change and biodiversity collapse (<xref ref-type="bibr" rid="B13845995">Pfenning-Butterworth et al. 2024</xref>). Although humanity itself is not physically limited by mountain topography, it nevertheless is confronted by analogous ecological boundaries, which also represent natural safeguards. The erosion of these natural safeguards threatens to turn Earth itself from a haven into a narrowing corridor of survival. Recognising mountains as both symbols and sentinels of change highlights the urgency of conservation and mitigation efforts. Protecting these ancient refuges is not only about preserving species, it is about preserving the conditions that have long made life possible. As the limits of ascent draw closer, both mountain endemics and humanity confront the same truth: survival depends on restoring the stability of our only refuge – Earth itself.</p>
    </sec>
  </body>
  <back>
    <sec sec-type="Conflicts of interest">
      <title>Conflicts of interest</title>
      <p>No conflict of interest to declare</p>
      <p>Disclaimer: This article is (co-)authored by any of the Editors-in-Chief, Managing Editors or their deputies in this journal.</p>
    </sec>
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  </back>
</article>
